*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

:root {
    --color-primary: #181d26;
    --color-primary-active: #0d1218;
    --color-canvas: #ffffff;
    --color-surface-soft: #f8fafc;
    --color-surface-strong: #e0e2e6;
    --color-surface-dark: #181d26;
    --color-surface-dark-elevated: #1d1f25;
    --color-hairline: #dddddd;
    --color-ink: #181d26;
    --color-body: #333840;
    --color-muted: #41454d;
    --color-on-primary: #ffffff;
    --color-link: #1b61c9;
    --color-link-active: #1a3866;
    --color-signature-coral: #aa2d00;
    --color-signature-forest: #0a2e0e;
    --color-signature-cream: #f5e9d4;
    --color-signature-peach: #fcab79;
    --color-signature-mint: #a8d8c4;
    --color-signature-yellow: #f4d35e;
    --rounded-xs: 2px;
    --rounded-sm: 6px;
    --rounded-md: 10px;
    --rounded-lg: 12px;
    --rounded-full: 9999px;
    --spacing-xxs: 4px;
    --spacing-xs: 8px;
    --spacing-sm: 12px;
    --spacing-md: 16px;
    --spacing-lg: 24px;
    --spacing-xl: 32px;
    --spacing-xxl: 48px;
    --spacing-section: 96px;
    --font-family: 'Inter', -apple-system, BlinkMacSystemFont, 'Segoe UI', Roboto, Oxygen, Ubuntu, Cantarell, 'Helvetica Neue', sans-serif;
    --max-width: 1280px;
}

html { font-size: 16px; scroll-behavior: smooth; }

body {
    font-family: var(--font-family);
    font-size: 14px;
    font-weight: 400;
    line-height: 1.5;
    color: var(--color-body);
    background: var(--color-canvas);
}

a { color: var(--color-link); text-decoration: none; }
a:hover { color: var(--color-link-active); }
img { display: block; max-width: 100%; height: auto; }
ul { list-style: none; }

.container {
    max-width: var(--max-width);
    margin: 0 auto;
    padding: 0 var(--spacing-xxl);
}

.site-nav {
    position: sticky;
    top: 0;
    z-index: 100;
    background: var(--color-canvas);
    border-bottom: 1px solid var(--color-hairline);
    height: 64px;
    display: flex;
    align-items: center;
}

.nav-inner {
    display: flex;
    align-items: center;
    justify-content: space-between;
    width: 100%;
    max-width: var(--max-width);
    margin: 0 auto;
    padding: 0 var(--spacing-xxl);
}

.nav-logo {
    font-size: 18px;
    font-weight: 500;
    color: var(--color-ink);
    letter-spacing: -0.3px;
}
.nav-logo span { color: var(--color-signature-coral); }

.nav-links {
    display: flex;
    align-items: center;
    gap: var(--spacing-xl);
}

.nav-links a {
    font-size: 14px;
    font-weight: 400;
    color: var(--color-body);
    transition: color 0.15s;
}
.nav-links a:hover { color: var(--color-ink); }
.nav-links a.active { color: var(--color-ink); font-weight: 500; }

.nav-hamburger {
    display: none;
    flex-direction: column;
    gap: 5px;
    cursor: pointer;
    background: none;
    border: none;
    padding: 4px;
}
.nav-hamburger span {
    display: block;
    width: 22px;
    height: 2px;
    background: var(--color-ink);
    border-radius: 2px;
    transition: all 0.2s;
}

.mobile-menu {
    display: none;
    position: fixed;
    top: 64px;
    left: 0;
    right: 0;
    bottom: 0;
    background: var(--color-canvas);
    z-index: 99;
    padding: var(--spacing-xl) var(--spacing-xxl);
    flex-direction: column;
    gap: var(--spacing-lg);
    overflow-y: auto;
}
.mobile-menu.open { display: flex; }
.mobile-menu a {
    font-size: 18px;
    font-weight: 400;
    color: var(--color-ink);
    padding: var(--spacing-sm) 0;
    border-bottom: 1px solid var(--color-hairline);
}

.hero-band {
    padding: var(--spacing-section) 0;
    background: var(--color-canvas);
}

.hero-content {
    max-width: 680px;
}

.hero-label {
    font-size: 13px;
    font-weight: 500;
    color: var(--color-muted);
    letter-spacing: 0.8px;
    text-transform: uppercase;
    margin-bottom: var(--spacing-md);
}

.hero-title {
    font-size: 40px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--color-ink);
    margin-bottom: var(--spacing-lg);
}

.hero-subtitle {
    font-size: 18px;
    font-weight: 400;
    line-height: 1.55;
    color: var(--color-body);
    margin-bottom: var(--spacing-xl);
    max-width: 560px;
}

.btn-primary {
    display: inline-block;
    background: var(--color-primary);
    color: var(--color-on-primary);
    font-size: 16px;
    font-weight: 500;
    line-height: 1.4;
    padding: 16px 24px;
    border-radius: var(--rounded-lg);
    border: none;
    cursor: pointer;
    text-decoration: none;
    transition: background 0.15s;
}
.btn-primary:hover { background: var(--color-primary-active); color: var(--color-on-primary); }

.btn-secondary {
    display: inline-block;
    background: var(--color-canvas);
    color: var(--color-ink);
    font-size: 16px;
    font-weight: 500;
    line-height: 1.4;
    padding: 16px 24px;
    border-radius: var(--rounded-lg);
    border: 1px solid var(--color-hairline);
    cursor: pointer;
    text-decoration: none;
    transition: border-color 0.15s;
}
.btn-secondary:hover { border-color: var(--color-ink); color: var(--color-ink); }

.btn-legal {
    display: inline-block;
    background: var(--color-link);
    color: var(--color-on-primary);
    font-size: 13.12px;
    font-weight: 600;
    padding: 12px 10px;
    border-radius: var(--rounded-xs);
    border: none;
    cursor: pointer;
    text-decoration: none;
}

.section-band {
    padding: var(--spacing-section) 0;
}

.section-title {
    font-size: 32px;
    font-weight: 400;
    line-height: 1.2;
    color: var(--color-ink);
    margin-bottom: var(--spacing-lg);
}

.section-subtitle {
    font-size: 16px;
    font-weight: 400;
    line-height: 1.55;
    color: var(--color-body);
    margin-bottom: var(--spacing-xl);
    max-width: 600px;
}

.articles-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: var(--spacing-lg);
}

.article-card {
    background: var(--color-canvas);
    border-radius: var(--rounded-md);
    overflow: hidden;
    border: 1px solid var(--color-hairline);
    transition: border-color 0.15s;
}
.article-card:hover { border-color: #b0b5be; }

.article-card-img {
    width: 100%;
    aspect-ratio: 16/9;
    object-fit: cover;
    border-radius: var(--rounded-md) var(--rounded-md) 0 0;
}

.article-card-body {
    padding: var(--spacing-md);
}

.article-card-tag {
    font-size: 11px;
    font-weight: 500;
    letter-spacing: 0.8px;
    text-transform: uppercase;
    color: var(--color-muted);
    margin-bottom: var(--spacing-xs);
}

.article-card-title {
    font-size: 18px;
    font-weight: 500;
    line-height: 1.4;
    color: var(--color-ink);
    margin-bottom: var(--spacing-xs);
}
.article-card-title a { color: inherit; }
.article-card-title a:hover { color: var(--color-link); }

.article-card-excerpt {
    font-size: 14px;
    color: var(--color-body);
    line-height: 1.5;
    margin-bottom: var(--spacing-md);
    display: -webkit-box;
    -webkit-line-clamp: 3;
    -webkit-box-orient: vertical;
    overflow: hidden;
}

.article-card-meta {
    font-size: 13px;
    color: var(--color-muted);
}

.signature-coral-card {
    background: var(--color-signature-coral);
    border-radius: var(--rounded-lg);
    padding: var(--spacing-xxl);
    color: var(--color-on-primary);
}

.signature-coral-card h2 {
    font-size: 32px;
    font-weight: 400;
    line-height: 1.2;
    margin-bottom: var(--spacing-lg);
    color: var(--color-on-primary);
}

.signature-coral-card p {
    font-size: 16px;
    line-height: 1.55;
    margin-bottom: var(--spacing-xl);
    max-width: 560px;
    opacity: 0.9;
}

.btn-on-dark {
    display: inline-block;
    background: var(--color-canvas);
    color: var(--color-ink);
    font-size: 16px;
    font-weight: 500;
    padding: 16px 24px;
    border-radius: var(--rounded-lg);
    border: none;
    cursor: pointer;
    text-decoration: none;
}
.btn-on-dark:hover { color: var(--color-ink); background: #f0f0f0; }

.dark-cta-card {
    background: var(--color-surface-dark);
    border-radius: var(--rounded-lg);
    padding: var(--spacing-xxl);
    color: var(--color-on-primary);
}

.dark-cta-card h2 {
    font-size: 32px;
    font-weight: 400;
    color: var(--color-on-primary);
    margin-bottom: var(--spacing-lg);
}

.dark-cta-card p {
    font-size: 16px;
    line-height: 1.55;
    margin-bottom: var(--spacing-xl);
    max-width: 560px;
    opacity: 0.85;
}

.cream-band {
    background: var(--color-signature-cream);
    padding: var(--spacing-section) 0;
}

.cream-band .section-title {
    color: var(--color-ink);
}

.species-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: var(--spacing-lg);
    margin-top: var(--spacing-xl);
}

.species-card {
    background: var(--color-canvas);
    border-radius: var(--rounded-md);
    padding: var(--spacing-lg);
    border: 1px solid var(--color-hairline);
}

.species-card img {
    width: 100%;
    aspect-ratio: 4/3;
    object-fit: cover;
    border-radius: var(--rounded-sm);
    margin-bottom: var(--spacing-md);
}

.species-card h3 {
    font-size: 18px;
    font-weight: 500;
    color: var(--color-ink);
    margin-bottom: var(--spacing-xs);
}

.species-card .species-latin {
    font-size: 13px;
    font-style: italic;
    color: var(--color-muted);
    margin-bottom: var(--spacing-sm);
}

.species-card p {
    font-size: 14px;
    line-height: 1.5;
    color: var(--color-body);
}

.cta-band-light {
    background: var(--color-surface-strong);
    border-radius: var(--rounded-lg);
    padding: var(--spacing-xxl);
    text-align: center;
    margin: var(--spacing-section) 0;
}

.cta-band-light h2 {
    font-size: 32px;
    font-weight: 400;
    color: var(--color-ink);
    margin-bottom: var(--spacing-md);
}

.cta-band-light p {
    font-size: 16px;
    color: var(--color-body);
    margin-bottom: var(--spacing-xl);
    max-width: 520px;
    margin-left: auto;
    margin-right: auto;
}

.contact-form-section {
    padding: var(--spacing-section) 0;
    background: var(--color-surface-soft);
}

.contact-form {
    max-width: 560px;
    margin: 0 auto;
}

.contact-form h2 {
    font-size: 32px;
    font-weight: 400;
    color: var(--color-ink);
    margin-bottom: var(--spacing-sm);
    text-align: center;
}

.contact-form .form-subtitle {
    font-size: 16px;
    color: var(--color-body);
    text-align: center;
    margin-bottom: var(--spacing-xl);
}

.form-row {
    margin-bottom: var(--spacing-md);
}

.form-row label {
    display: block;
    font-size: 14px;
    font-weight: 500;
    color: var(--color-ink);
    margin-bottom: var(--spacing-xs);
}

.form-row input,
.form-row textarea {
    width: 100%;
    background: var(--color-canvas);
    color: var(--color-ink);
    font-size: 14px;
    font-family: var(--font-family);
    padding: 12px 16px;
    height: 44px;
    border-radius: var(--rounded-sm);
    border: 1px solid var(--color-hairline);
    outline: none;
    transition: border-color 0.15s;
}

.form-row textarea {
    height: auto;
    min-height: 100px;
    resize: vertical;
}

.form-row input:focus,
.form-row textarea:focus {
    border-color: #458fff;
}

.form-submit {
    width: 100%;
    padding: 16px;
    font-size: 16px;
    font-weight: 500;
    margin-top: var(--spacing-sm);
}

.site-footer {
    background: var(--color-canvas);
    border-top: 1px solid var(--color-hairline);
    padding: var(--spacing-section) 0 var(--spacing-xl);
}

.footer-inner {
    display: grid;
    grid-template-columns: 2fr 1fr 1fr 1fr;
    gap: var(--spacing-xl);
    margin-bottom: var(--spacing-xxl);
}

.footer-brand .nav-logo {
    display: inline-block;
    margin-bottom: var(--spacing-md);
    font-size: 18px;
}

.footer-brand p {
    font-size: 14px;
    color: var(--color-muted);
    line-height: 1.6;
    max-width: 280px;
    margin-bottom: var(--spacing-md);
}

.footer-contact {
    font-size: 13px;
    color: var(--color-muted);
    line-height: 2;
}

.footer-col h4 {
    font-size: 14px;
    font-weight: 500;
    color: var(--color-ink);
    margin-bottom: var(--spacing-md);
}

.footer-col ul li {
    margin-bottom: var(--spacing-xs);
}

.footer-col ul li a {
    font-size: 14px;
    color: var(--color-muted);
    transition: color 0.15s;
}
.footer-col ul li a:hover { color: var(--color-ink); }

.footer-bottom {
    display: flex;
    align-items: center;
    justify-content: space-between;
    padding-top: var(--spacing-xl);
    border-top: 1px solid var(--color-hairline);
    font-size: 13px;
    color: var(--color-muted);
}

.footer-bottom-links {
    display: flex;
    gap: var(--spacing-lg);
}
.footer-bottom-links a {
    color: var(--color-muted);
    font-size: 13px;
}
.footer-bottom-links a:hover { color: var(--color-ink); }

.cookie-banner {
    display: none;
    position: fixed;
    bottom: 0;
    left: 0;
    right: 0;
    background: var(--color-surface-dark);
    color: var(--color-on-primary);
    padding: var(--spacing-lg) var(--spacing-xxl);
    z-index: 200;
    border-top: 1px solid #2d3340;
}

.cookie-banner.visible { display: flex; }

.cookie-inner {
    display: flex;
    align-items: center;
    justify-content: space-between;
    gap: var(--spacing-xl);
    max-width: var(--max-width);
    margin: 0 auto;
    width: 100%;
}

.cookie-text {
    font-size: 14px;
    line-height: 1.6;
    opacity: 0.85;
    flex: 1;
}

.cookie-text a {
    color: #7eb8ff;
    text-decoration: underline;
}

.cookie-buttons {
    display: flex;
    gap: var(--spacing-sm);
    flex-shrink: 0;
}

.page-hero {
    padding: var(--spacing-section) 0 var(--spacing-xxl);
    border-bottom: 1px solid var(--color-hairline);
    margin-bottom: var(--spacing-section);
}

.page-hero h1 {
    font-size: 40px;
    font-weight: 400;
    color: var(--color-ink);
    margin-bottom: var(--spacing-md);
}

.page-hero p {
    font-size: 18px;
    color: var(--color-body);
    max-width: 600px;
}

.page-meta {
    font-size: 13px;
    color: var(--color-muted);
    margin-top: var(--spacing-sm);
}

.prose {
    max-width: 720px;
    margin: 0 auto;
}

.prose h1 {
    font-size: 40px;
    font-weight: 400;
    color: var(--color-ink);
    margin-bottom: var(--spacing-lg);
    line-height: 1.2;
}

.prose h2 {
    font-size: 28px;
    font-weight: 400;
    color: var(--color-ink);
    margin-top: var(--spacing-xxl);
    margin-bottom: var(--spacing-md);
    line-height: 1.3;
}

.prose h3 {
    font-size: 20px;
    font-weight: 500;
    color: var(--color-ink);
    margin-top: var(--spacing-xl);
    margin-bottom: var(--spacing-sm);
}

.prose p {
    font-size: 16px;
    line-height: 1.7;
    color: var(--color-body);
    margin-bottom: var(--spacing-lg);
}

.prose ul, .prose ol {
    margin-bottom: var(--spacing-lg);
    padding-left: var(--spacing-xl);
}

.prose li {
    font-size: 16px;
    line-height: 1.7;
    color: var(--color-body);
    margin-bottom: var(--spacing-xs);
    list-style: disc;
}

.prose ol li { list-style: decimal; }

.prose a { color: var(--color-link); }
.prose a:hover { color: var(--color-link-active); }

.prose img {
    width: 100%;
    border-radius: var(--rounded-md);
    margin: var(--spacing-xl) 0;
}

.prose .article-img-caption {
    font-size: 13px;
    color: var(--color-muted);
    text-align: center;
    margin-top: calc(-1 * var(--spacing-md));
    margin-bottom: var(--spacing-xl);
}

.article-layout {
    display: grid;
    grid-template-columns: 1fr 300px;
    gap: var(--spacing-xxl);
    align-items: start;
    padding: var(--spacing-section) 0;
}

.article-sidebar {
    position: sticky;
    top: 84px;
}

.sidebar-card {
    background: var(--color-surface-soft);
    border-radius: var(--rounded-md);
    padding: var(--spacing-lg);
    margin-bottom: var(--spacing-lg);
    border: 1px solid var(--color-hairline);
}

.sidebar-card h4 {
    font-size: 16px;
    font-weight: 500;
    color: var(--color-ink);
    margin-bottom: var(--spacing-md);
}

.sidebar-links li {
    margin-bottom: var(--spacing-xs);
}

.sidebar-links li a {
    font-size: 14px;
    color: var(--color-link);
}

.about-grid {
    display: grid;
    grid-template-columns: 1fr 1fr;
    gap: var(--spacing-xxl);
    align-items: center;
    padding: var(--spacing-section) 0;
}

.about-img {
    width: 100%;
    aspect-ratio: 4/3;
    object-fit: cover;
    border-radius: var(--rounded-lg);
}

.about-text h2 {
    font-size: 32px;
    font-weight: 400;
    color: var(--color-ink);
    margin-bottom: var(--spacing-lg);
}

.about-text p {
    font-size: 16px;
    line-height: 1.7;
    color: var(--color-body);
    margin-bottom: var(--spacing-md);
}

.values-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: var(--spacing-lg);
    margin-top: var(--spacing-xl);
}

.value-card {
    background: var(--color-surface-soft);
    border-radius: var(--rounded-md);
    padding: var(--spacing-lg);
    border: 1px solid var(--color-hairline);
}

.value-card h3 {
    font-size: 18px;
    font-weight: 500;
    color: var(--color-ink);
    margin-bottom: var(--spacing-sm);
}

.value-card p {
    font-size: 14px;
    line-height: 1.6;
    color: var(--color-body);
}

.privacy-content,
.terms-content {
    max-width: 800px;
    margin: 0 auto;
    padding: var(--spacing-section) 0;
}

.privacy-content h2,
.terms-content h2 {
    font-size: 24px;
    font-weight: 400;
    color: var(--color-ink);
    margin-top: var(--spacing-xxl);
    margin-bottom: var(--spacing-md);
}

.privacy-content p,
.terms-content p {
    font-size: 16px;
    line-height: 1.7;
    color: var(--color-body);
    margin-bottom: var(--spacing-md);
}

.privacy-content ul,
.terms-content ul {
    padding-left: var(--spacing-xl);
    margin-bottom: var(--spacing-md);
}

.privacy-content li,
.terms-content li {
    font-size: 16px;
    line-height: 1.7;
    color: var(--color-body);
    list-style: disc;
    margin-bottom: var(--spacing-xs);
}

.disclaimer-box {
    background: var(--color-signature-cream);
    border-radius: var(--rounded-md);
    padding: var(--spacing-lg);
    margin-top: var(--spacing-xl);
    font-size: 14px;
    color: var(--color-body);
    line-height: 1.6;
}

.breadcrumb {
    font-size: 13px;
    color: var(--color-muted);
    margin-bottom: var(--spacing-lg);
}

.breadcrumb a {
    color: var(--color-muted);
}
.breadcrumb a:hover { color: var(--color-ink); }
.breadcrumb span { margin: 0 6px; }

@media (max-width: 1024px) {
    .articles-grid { grid-template-columns: repeat(2, 1fr); }
    .species-grid { grid-template-columns: repeat(2, 1fr); }
    .footer-inner { grid-template-columns: 1fr 1fr; }
    .article-layout { grid-template-columns: 1fr; }
    .article-sidebar { position: static; }
    .values-grid { grid-template-columns: 1fr 1fr; }
}

@media (max-width: 768px) {
    :root { --spacing-section: 64px; --spacing-xxl: 24px; }
    .container { padding: 0 var(--spacing-md); }
    .nav-inner { padding: 0 var(--spacing-md); }
    .nav-links { display: none; }
    .nav-hamburger { display: flex; }
    .hero-title { font-size: 28px; }
    .hero-subtitle { font-size: 16px; }
    .articles-grid { grid-template-columns: 1fr; }
    .species-grid { grid-template-columns: 1fr; }
    .footer-inner { grid-template-columns: 1fr; }
    .footer-bottom { flex-direction: column; gap: var(--spacing-md); text-align: center; }
    .about-grid { grid-template-columns: 1fr; }
    .values-grid { grid-template-columns: 1fr; }
    .cookie-inner { flex-direction: column; }
    .signature-coral-card { padding: var(--spacing-xl); }
    .dark-cta-card { padding: var(--spacing-xl); }
    .cta-band-light { padding: var(--spacing-xl); }
    .section-title { font-size: 24px; }
    .page-hero h1 { font-size: 28px; }
    .prose h1 { font-size: 28px; }
    .prose h2 { font-size: 22px; }
}